RT not sending email - No recipients found

I have just setup my new web server and successfully installed RT as per
instructions. Everything seems to work fine, I can email in a ticket and
get an autoreply from RT, however when I attempt to add a comment, or
reply within the ticket no email is sent to the requestor. I have turned
on debugging and found these entries in the rt.log file.

rt-3.4.2-3-104-5.12.837616257616@adastral.ucl.ac.uk #3/104 - Scrip 5
(/usr/local/rt/lib/RT/Action/SendEmail.pm:235)
rt-3.4.2-3-104-5.12.837616257616@adastral.ucl.ac.uk No recipients
found. Not sending. (/usr/local/rt/lib/RT/Action/SendEmail.pm:247)

I am fairly unfamiliar with RT, but have read most of the documentation
but can’t find the answer, can anyone help?

OS : RHEL4
RT version 3.4.2

Many thanks,

Neil.

Neil Marjoram
Systems Manager
Adastral Park Campus
University College London
Ross Building
Adastral Park
Martlesham Heath
Ipswich - Suffolk
IP5 3RE

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